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新疆准噶尔盆地北缘中中新世猬类化石
ERINACEIDAE FROM THE MIDDLE MIOCENE OF NORTH JUNGGAR BASIN,XINJIANG UYGUR AUTONOMOUS REGION,CHINA
【Author】 BI Shundong;WU Wenyu;YE Jie;MENG Jin;Insitute of Vertebrate Paleontology and Paleoanthropology,Chinese Academy of Sciences;Department of Biology,University of Massachusetts;
【机构】 中国科学院古脊椎动物与古人类研究所; 美国马萨诸塞州州立大学生物系;
【摘要】 记述了新疆准噶尔盆地北缘乌伦古河南岸夺勒布勒津中中新世的两种猬类化石Mioechinus?cf.M.gobiensis和Schizogalerix duolebulejinensis sp.nov.。Schizogalerix在我国是首次发现,S.duolebulejinensis sp.nov.在形态上与已知的欧洲、土耳其和北非的Schizogalerix属的各个支系均有所不同,应为一个独立的支系。
【Abstract】 During four field seasons of 1995~1998,insectivores were collected from the late Oligocene through the middle Miocene at Chibaerwoyi,Duolebulejin,Tieershihabahe of north Junggar Basin,Xinjiang Uygur Autonomous Region.Most of them belong to Erinaceidae,of which two forms from the early Miocene of Chibaerwoyi have been described(Bi,1999,In press).The present paper describes the erinaceids from the middle Miocene of Duolebulejin.Mioechinus? cf.M.gobiensis Qiu,1996(P1.I,1,3)Nine isolated teeth(3 M1,3 M2,1 p4,1 m1,lm2.IVPP V11815.1~9) from lower part of Halamagai Formation of the early middle Miocene,Duolebulejin,are described.The Junggar form is similar to Mioechinus? gobiensis from Tunggur[4] in size and dental morphology,but differs from the latter in having a deeper groove on the lingual root of M1~2,stronger parastyle on M2,and more developed paralophid on p4.Mioechinus resembles Amphechinus in dental morphology.It is difficult to distinguish them without i1[4].Following Qiu[4],the Junggar form is assigned to Miochinus with question mark.Schizogalerix duolebulejinensis sp.nov.(P1.I,2,4~5)Holotype IVPP V11816.1(P1.I,5 a~b),fragmentary right mandible with p4~m2 and the anterior root of m3.Type locality Duolebulejin,Fuhai County,Xinjiang Uygur Autonomous Region.Horizon and age upper sands of uppermost Suosuoquan Formation,early middle Miocene.Referred specimen IVPP V11816.2,1 right M1,from upper sands of uppermost Suosuoquan Formation.IVPP V11817.1~4,4 isolated teeth(1 left P4,1 left M1,2 right m1),from basal part of Halamagai Formation.Measurements See the Chinese text Diagnosis Large in size;anterolingual-posterolabially elongated P4 and M1;M1 with split mesostyles connected at the base,stronger parastyle,and the absence of the posterior arm of hypocone;smaller p4 relative to lower molars;m1~2 with the entoconid connected to the posterior cingulum and the posterior arm of hypoconid postero-interially,developed metastylid on m1~2.Etymology Duolebulejin,prefix,indicate the locality where the species was found for the first time.Description The holotype V11816.1(Pl.I,5 ab):the mandible is 3.60 mm high at the level of the posterior root of p4 and 3.55 mm at the level of the anterior root of m2 on the labial side.The mandible is broken in front of p4 and behind m3.The absence of the mental foramen on the remainder ramal suggests that it be located in front of p4.p4 is two rooted.The occlusal surface is triangle in shape.The metaconid is slightly lower than the protoconid.The paraconid is rather low,which is connected to the protoconid by weak paralophid.The posterior cingulum is developed with a tubercle in the middle.The labial cingulum is weak.The trigonid of m1 is short.There are striped grains on the labial surface of the trigonid.The protoconid and metaconid are the same high.The paraconid is slightly lower than the protoconid and connected to the latter by the developed parapholid.The metastylid is developed and ridge-like,which is separated from the entoconid by a deep notch.The talonid is broad.The entoconid meets posteriorly the posterior cingulum and the posterior arm of the hypoconid around the posterointernal corner of the tooth.A shallow notch is present in the middle of the posterior arm of the hypoconid.The labial cingulum is short but developed,being separated from the anterior cingulum at the base of the protocone.Tworooted.The m2 resembles the m1,but differs from the latter in small size,the shorter trigonid,and the bladelike paraconid,which connects with the paralophid.P4(Pl.I,4) is three-rooted,and anterolingual-posterolabially elonglated.The high paracone is the largest cusp.The posteriorcrista runs from its top backward,then bends halfway sharply to the posterolabial corner of the tooth.The paracone is separated from the lingual cusps by a wide valley.The protocone is higher than the hypocone and anteroligual to the latter.A shallow valley is between these two cusps.The parastyle is distinct.The posterior cingulum is developed.M1(Pl.I,2) is large and anterolingual-posterolabially elonglated.The paracone and metacone are pyramid-shaped.The paracone is slightly lower than the latter.The protocone is the largest cusp,the anterior arm of which runs parallel to the anterior border of the tooth and connects with the protoconule;the protocone is posteriorly connected with the hypocone by an indistinct ridge,but separated from the metaconule by a deep valley.The hypocone is cone-shaped.The parastyle and metastyle are distinct.The mesostyle is split,but slightly joined at the base.The metaconule is crescent-shaped.Its anterior arm ends against the base of the metacone and its long and oblique posterior arm ends against the base of the metastyle.The protoconule is D-shaped.A well developed cingulum is present on the anterior and posterior border of the tooth respectively.The labial cingulum is very weak.Three-rooted.Remarks The Schizogalerix described here comes from two different not far apart levels at Duolebulejin.They are identical in size and morphology.Moreover,the components of associated faunas are nearly the same.Therefore,the whole sample is treated as representing one species.Judging from the morphology of the M1:longer than wide,the split mesostyle,the posterior arm of the protocone separated from the metaconule,and the strong mandible,the Junggar form undoubtedly belongs to the genus Schizogalerix.So far,at least nine species have been described in the genus.The Junggar form is a large species of Schizogalerix.It is much larger than all the known species of this genus except for S.nov.sp.Amama II Engesser,1980.The Junggar form still differs from all known species in the two distinct features:1)The well-developed metastylid on m1 and m2.2) The posterior arm of the entoconid,the posterior cingulum and the posterior arm of the hypoconid join around the posterointernal corner of the m1 and m2.It is assigned to the new species,Schizogalerix duolebulejinensis.The Junggar form is so far the first record of the genus in China and it is the easternmost species of the genus.
- 【会议录名称】 第七届中国古脊椎动物学学术年会论文集
- 【会议名称】第七届中国古脊椎动物学学术年会
- 【会议时间】1999-04-25
- 【会议地点】中国云南玉溪
- 【分类号】Q915.2
- 【主办单位】中国科学院古脊椎动物与古人类研究所、云南省玉溪市政府文化局