In the pollen germinated for more than 4 hours,many cytoplasmic fibrils can be observ- ed under optical microscope.These fibrils which form a network are always in motion,being rigid and flexible to a certain extent,defined as moving cytoplasmic fibril network in the present paper.The network fibrils are different from cytoplasmic streaming,because they not only move vertically but also horizontally.Furthermore,they restrict the transportation of cytoplasmic particles.Electron micrographs show that the fibr...