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椭圆蕉羊齿羽轴解剖及蕉羊齿属的系统分类位置

ANATOMICAL STUDY OF THE RACHIS OF COMPSOPTERIS ELLIPTICA AND THE SYSTEMATIC POSITION OF COMPSOPTERIS

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【作者】 郭英廷田宝霖韩德馨

【Author】 Guo Ying-ting Tian Bao-lin Han De-xin (Beiiing Graduate School, China University of Mining and Technology, Beijing 100083)

【机构】 中国矿业大学北京研究生部

【摘要】 描述了产自贵州省盘县矿区晚二叠世含煤地层中的椭圆蕉羊齿(Compsopteris ellipticecx Yang et Chen 化石,其主要特征如下:羽状复叶,羽轴具背腹性;羽轴维管束2环或2环半,腹面凹陷;维管束外韧式,木质部内始式,原生木质部管胞螺纹加厚,后生木质部管胞梯纹加厚;各维管束环之间具厚壁细胞带;皮层分内外2层,外层为薄壁细胞,内层为厚壁细胞。据上述特征,作者认为蕉羊齿属(Compsopteris)(至少华南地区的标本)并非为种子蕨植物,而是真蕨中的观音座莲目(Marattiales)植物。

【Abstract】 The anatomical characters of the rachis of Compsopteris elliplica ex Yang et Chen are des- cribed from the calc-petrified specimens of Lhe Lare Permian of Panxing, Guizhou, China. The main characters of its rachis are as follows (Plate I, 1—4; 6—9): The vascular bundles of 2— 2.5 cycles and ectophloic type. Phloem, consisting of 1—2 layers of. cells. Protoxylem, compo- sed of several small tracheids which are less than 20μm in diameter. Metaxylem, consisting of 2--- 3 layers of. tracheids, about 30μm in diameter, scalariform thickenning. The sclerenchyma zone lies between the vascular bundles, its cell, small, 20—25 μm in diameter, some containing brown substances. The cortex could be divided into two zones: outer zone, consisting of thin-walled cells, and inner zone, thicken-walled cells, with secretory cavities. Epidermis, one layer, rectan- gular, some with contents. Compsopteris sp. (Plate l,5) 3—4 cycles vascular bundles, the same as C. elli- ptica in many respects; only larger than C. ellipptica in diameter. It may represent the base part of the rachis or larger rachis. Based on the shape of vascular bundles and the structure of xylem and phloem, Compsop- ,eris is very similar to Angiopteris, Danaea etc., which all belong to the Marattiales. In addi’ion, Huang et al. (1989) found that the sporangia of Compsopteris is similar to that of Danaeltes, which belongs to the Marattiales. Therefore, Compsopteris does not belong to the seed fern, but the Marattiales.

  • 【文献出处】 Journal of Integrative Plant Biology ,植物学报(英文版) , 编辑部邮箱 ,1992年08期
  • 【被引频次】2
  • 【下载频次】52
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