【摘要】 知母(Anemarrhena asphodeloides)的雄配子体发育有三个特点:1.除高尔基小泡直接参与胼胝质壁物质的形成外,其他小泡的活动可能与脂体物质的沉积有关。2.生殖细胞形成时,脂体分布发生明显的极性现象,它们沿着营养细胞与生殖细胞两层交界膜排列,形成所谓脂体冠。3.在小孢子阶段,细胞质中具有丰富的质体,但在第一次有丝分裂之后,全部集中在营养细胞中,而生殖细胞则完全缺乏质体。更多还原

【Abstract】 Anemarrhena asphodeloides is a monotypic genus of Liliaceae,endemic to China and Korea.This genus is characterized by possessing three stamens.From development of male gametophyte,three features of the species are noteworthy.(1) During meiosis of the micros- pore mother cells,the Golgi vesicles are immediately incorporated into the formation of the material of callose wall;The latter lying at the oute,tangential is about 4 μm in thickness during formation of the tetrad.In the outer tangential callose wall there are certain cytoplas- mic canals,which are about 0.6 to 1 μm in diameter.During the development of‘pollen grains,there are a number of other vesicles dispeising in the cytoplasm of the microspores.The activity of these vesicles seems to be involved in accumulation and formation of lipid bodies. But the above vesicles,which were derivxed from Golgi or endoplasmic reticulum,have not been known in this genus.(2) By two-celled stage of pollen grains,the unequal distribution of lipid bodies is very prominent,and they are singular in being placed on the boundary be- tween the plasmalemma of vegetative and generative ceils.While the generative cell is de- Iached from the intine of pollen grain,the generative cell is surrounded by the lipid bodies which had been called the corona of them.By the observation of TEM, these lipid bodies come from the cytoplasm of vegetative cell and did not remain a constant surroundint layer. Towards the stage of pollen maturation,the lipid bodies-lying oppositely to the nucleus Of ve- gerative cell were gradually dispersed in the cytoplasm.Their function is unknown but the observation shows that some of them move to the plasmalemma of the pollen grain.(3)An important feature of the mature pollen grain in Anemarrhena is that the generative cell does not contain plastids during polle development.On the basis of cytological mechanisms of the plastid inheritance,Hagemann (1983) has classified the angiosperms into four groups of species,of which the Lycopersicnm type,Solanllm type,and Triticum type belong to the mode of a uniparental maternal inheritance of plastids;while the Pelargonium type represents the mode of biparental inheritance of plastids.Our studies have confirmod that the mode of piastid inheritance in dnemarrhena asphodeloides is similar to Gasteria verrucosa,both show the same mode of plastid inheritance of Lycopersicom type.更多还原